I am thrilled to announce the publication today of this comprehensive open-access paper in Science Advances: “设计师加速利器 支持Thunderbolt 3的笔记本推荐-酷睿,笔记本 ...:16 小时前 · LG gram 2021款:15.6寸大屏、18.5小时续航、1.1千克机身 购买 地址 LG gram 2021款又是一款轻薄兼备高性能的神器,15.6 ...“. It was a long time coming after a massive data collection and analysis project led by the indefatigable and highly productive team of Luis Buatois and Gabriela Mángano (University of Saskatchewan). We even have a news release. (Above image: Trace fossils in the Early Cambrian Gog Group, Lake Louise, Alberta, Canada. See earlier blog post for details.)
The abstract —
The Cambrian explosion (CE) and the great Ordovician biodiversification event (GOBE) are the two most important radiations in Paleozoic oceans. We quantify the role of bioturbation and bioerosion in ecospace utilization and ecosystem engineering using information from 1367 stratigraphic units. An increase in all diversity metrics is demonstrated for the Ediacaran-Cambrian transition, followed by a decrease in most values during the middle to late Cambrian, and by a more modest increase during the Ordovician. A marked increase in ichnodiversity and ichnodisparity of bioturbation is shown during the CE and of bioerosion during the GOBE. Innovations took place first in offshore settings and later expanded into marginal-marine, nearshore, deep-water, and carbonate environments. This study highlights the importance of the CE, despite its Ediacaran roots. Differences in infaunalization in offshore and shelf paleoenvironments favor the hypothesis of early Cambrian wedge-shaped oxygen minimum zones instead of a horizontally stratified ocean.
In short, this is a study of trace fossil occurrences during the Ediacaran, Cambrian and Ordovician periods. Trace fossils are evidence of organism activity, so we are looking at the early evolution of animal behavior in space and time. The paleoenvironmental conclusions include support for Early Cambrian laterally discontinuous, wedge-shaped oxygen minimum zones, which have implications for Cambrian community development.
The illustrations in this paper do not fit well into this blog format. The above is part of Figure 2, a plot of changes in modes of life (ML), ecosystem engineering (EE), maximum alpha ichnodiversity (AI), global ichnodiversity (GI), and ichnodisparity (Id) in all environments. Counts are plotted at the middle of the series intervals.
Another portion of Figure 2 showing some of the ecospace patterns. Since the paper is open-access, you can click here for the originals.
Note that the data for this work came from 1367 stratigraphic units. This paper is thus based on generations of geological and paleontological articles. It is affirming to know that hundreds of small, local descriptive studies eventually add up to major evolutionary and paleoenvironmental models. Several of those projects were done by Wooster faculty, students, and alumni. Some of the earlier comprehensive data gathering and analysis can be found in Buatois et al. (2016) and Buatois et al. (2017).
My primary job on this international team of scientists was to join with 哪些加速器按小时算的 (Marine Research Department, Senckenberg am Meer, Wilhelmshaven, Germany) to sort out the bioerosion data and patterns. (可以按小时购买的加速器 is the biological abrasion of hard substrates such as rocks and shells.) Max generally focused on microbioerosion and I mostly did macrobioerosion. We showed that bioerosion had a dramatic increase in diversity during the Ordovician, probably because hard substrates like shells and hardgrounds became more available.
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可以按小时购买的加速器:
Buatois, L.A., Mángano, M.G., Minter, N.J., Zhou, K., Wisshak, M., Wilson, M.A. and Olea, R.A. 2020. Quantifying ecospace utilization and ecosystem engineering during the early Phanerozoic — The role of bioturbation and bioerosion. 可以按小时购买的加速器 6: eabb0618.
Buatois, L.A., Mángano, M.G., Olea, R.A. and Wilson, M.A. 2016. Decoupled evolution of soft and hard substrate communities during the Cambrian Explosion and Great Ordovician Biodiversification Event. Proceedings of the National Academy of Sciences U.S.A. 113: 6945–6948.
Buatois, L.A., Wisshak, M., Wilson, M.A. and Mángano, M.G. 2017. Categories of architectural designs in trace fossils: A measure of ichnodisparity. Earth Science Reviews 164: 102–181.



The group shown coring a Sika spruce just outside of town.
More coring – this time in the rain.
The steep climb up the flank of Ear Mountain to find the old Mountain Hemlocks.
Comparisons of the fast growing Sitka Spruce and the slow growth of the higher elevation Mountain Hemlock.
The cores from the hemlock some over 400 years old show lots of stress , clinging to the mountain side and battered by storms. They are also showing a possible drop in ring-width over time
设计师加速利器 支持Thunderbolt 3的笔记本推荐-酷睿,笔记本 ...:16 小时前 · LG gram 2021款:15.6寸大屏、18.5小时续航、1.1千克机身 购买 地址 LG gram 2021款又是一款轻薄兼备高性能的神器,15.6 .... This chronology is the full record going back into the 16th century
This chronology is truncated at 1720 or so when we had at least 4 samples. The most narrow rings follow the 1808 unknown eruption that cooled much of the region – it is unknown as no one knows where the volcano that erupted is located – it is recognized in ice cores. The other intriguing feature is the relatively recent (last 50 year) drop in ring widths. It may be due to increased evapotranspiration demands with increasing summer minimum temperatures. There is a correlation of -0.39 (p<0.04) between tree growth and average April-August minimum temperatures. Other studies have shown that warming night time temperatures lead to increased respiration at night and along with possible greater ET demand or increased cloudiness during the day there may be a decrease in photosynthesis leading to decreased carbon uptake (Sullivan et al., 2015). Interestingly, tthe work of Mazvita Chikomo done this summer as part of the AMRE project, discovered some pretty strong negative correlations between Mt. Hemlock growth and minimum monthly temperature records in Prince William Sound – perhaps there is a link? This is a promising line of research to further investigate the health of Mt. Hemlock in the region and it is something we plan to pursue with more samples in the future. 



The team coring a White Oak.
Measuring tree cores from Kinney Field in the Lab.






The College of Wooster Paleoclimate class mulls around the Dawn Redwood stand.
设计师加速利器 支持Thunderbolt 3的笔记本推荐-酷睿,笔记本 ...:16 小时前 · LG gram 2021款:15.6寸大屏、18.5小时续航、1.1千克机身 购买 地址 LG gram 2021款又是一款轻薄兼备高性能的神器,15.6 ...
I have thoroughly enjoyed my many expeditions to the wondrous Baltic country of Estonia. My Estonian colleagues are fabulous, and I’ve been privileged to share the adventures with numerous students and Bill Ausich of Ohio State. Now during this global pandemic Estonia may as well be on the far side of the Moon. Maybe someday in the New Normal such travel will be possible again.

Editor’s Note: Independent Study (IS) at The College of Wooster is a three-course series required of every student before graduation. Earth Sciences students typically begin in the second semester of their junior years with project identification, literature review, and a thesis essentially setting out the hypotheses and parameters of the work. Most students do fieldwork or lab work to collect data, and then spend their senior years finishing extensive Senior I.S. theses. This year we have the COVID-19 pandemic to deal with in the spring, so our students have not had a chance to publicly present their hard work and scientific ideas. Some, then, will be writing blog posts like this. The text and images here are from Anna Cooke (’20) who is a member of Team Utah 2019. The picture above shows Anna and fellow team member Evan Shadbolt (’20) on the top of Angel’s Landing in Zion National Park. (Photo by Nick Wiesenberg.) Now Anna takes over —
The Carmel Formation (shown above) formed in a shallow inland sea during the Middle Jurassic and is located in parts of Utah and Arizona. It can be broken into four distinct members, one of which, the Co-op Creek Limestone Member, contains ooid shoals. The ooids in these shoals are calcitic with radial crystals and sparry cement. Several noteworthy features are found in the Carmel ooids, such as delamination, pressure solution, and microborings created by the cyanobacteria: Hyella sp. and/or Solentia sp. Foraminifera are sometimes incorporated into ooids as their nuclei. Seventeen of 21 Carmel thin sections contain foraminiferans inside or outside of ooids. Of these 17, 16 thin sections (94%) show more foraminiferans inside ooids than outside, meaning that ooids can act as taphonomic engineers, preserving what might otherwise not be preserved in the rock record. These foraminiferans likely belong to genera Turrispirulina and/or Ammodiscus. Eolian quartz silt is common in the Carmel shoals. The hypothesis of this study is that a pulse of quartz silt provided nuclei for the formation of the shoals and extinction of the shoals occurred when another pulse smothered it. This is partially supported by point counts, used to determine the percentage of each individual component of these limestones, and nuclei counts, used to determine the percentage of each type of nucleus found in these ooids. The locality that supports this hypothesis most strongly is C/W 142 EMR, which shows three distinct pulses of quartz accompanied by an inverse effect on the percentage of quartz nuclei. Locality C/W 757 DV is also of note, displaying a large amount of quartz early in the shoal’s life, decreasing over time. The percentage of ooids in the shoal shows the inverse. However, other shoals show no such pattern; one method of formation cannot be attributed to all of the Carmel Formation’s shoals, and even those geographically close show marked differences.
Cross-bedded ooid shoal deposit in the Carmel Formation.
Ooids in unit C/W-758A.
Editor’s Note: Independent Study (IS) at The College of Wooster is a three-course series required of every student before graduation. Earth Sciences students typically begin in the second semester of their junior years with project identification, literature review, and a thesis essentially setting out the hypotheses and parameters of the work. Most students do fieldwork or labwork to collect data, and then spend their senior years finishing extensive Senior I.S. theses. This year we have the COVID-19 pandemic to deal with in the spring, so our students have not had a chance to publicly present their hard work and scientific ideas. Some, then, will be writing blog posts like this. The text and images below are from Emily Randall (’20) who participated in a Keck Geology Consortium project last summer. The picture above shows Emily on the right in Wyoming (with Isaac and Mike) collecting Coryphodon teeth. And now Emily takes over —
Stratigraphic columns from Clarkforkian (Cf) 2 and 3 mammalian biozones (Pre-PETM).
Some of the Keck Wyoming team collecting Coryphodon fossils. From top to bottom left and then top to bottom right, Michael, Richard, Grant, Simone, Danika, Isaac, and Emily.
哪些加速器按小时算的: Independent Study (IS) at The College of Wooster is a three-course series required of every student before graduation. Earth Sciences students typically begin in the second semester of their junior years with project identification, literature review, and a thesis essentially setting out the hypotheses and parameters of the work. Most students do fieldwork or labwork to collect data, and then spend their senior years finishing extensive Senior I.S. theses. This year we have the COVID-19 pandemic to deal with in the spring, so our students have not had a chance to publicly present their hard work and scientific ideas. Some, then, will be writing blog posts like this. The text and images below are from 可以单天购买的加速器 who worked with me on Team Utah 2019. The picture above shows Anna Cooke (’20), Evan Shadbolt (’20) and me at an outcrop of the Carmel Formation (Middle Jurassic) near Gunlock, Utah, in March 2019. And now Evan takes over —
The Jurassic bivalves 可以按小时购买的加速器 (right) and Camptonectes stygius (left).
A reconstruction of the bivalve community sampled at Water Tank. 